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1977-1982 - MS in Biochemistry, Department of Biochemistry, Moscow Veterinary-Medicine Academy, Moscow, Russia
1982-1986 - Diploma in Molecular Biology and Genetic Engineering, Special Higher Qualification Courses, Branch Division of Lomonosov State University of Moscow, Pushchino, Russia
1986-1991 - PhD in Molecular Biology, Engelhardt Institute of Molecular Biology, Russian Academy of Sciences, Moscow, Russia
1991-1995 - Research Scientist, Engelhardt Institute of Molecular Biology, Russian Academy of Sciences, Moscow, Russia
1995-1996 - Research Fellow, University of St. Andrews, St. Andrews, Scotland
1996-1997 - Research Scientist, Engelhardt Institute of Molecular Biology, Russian Academy of Sciences, Moscow, Russia
1997-2000 - Research Fellow, Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, MI
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